Descriptions based upon the Clusters (Groups) identified by MCA.

(Character numbers bracketed, as described in the measurements spread sheet Cyperus papyrus Measurements:Legend & Chars;  ave. = average)

Group 1

Stems usually bluntly trigonous, occasionally sharply triquetrous.  Rays R1 crowded, rarely 25-40, generally 40+ (46), ave. 65 - 342mm long (5).  Bracts B1 erect to spreading (53), length variable ave. (6.3 -) 40 - 98mm (2); prophylls P1 ave. 19 - 37mm (9) long.  Spikes ave. (1 -) 3 - 4 (-5) (48-50), ave. length 4 - 10.8mm (11); prophylls P2 ave. 0.5 - 3.8mm (21);  longest rachis R2 ave. 2-9 (- 22)mm (51),  sessile rachis’ R2 ave. 0 - 1, rachis’ R2 <10mm (0.3) - 2.0 - 4.3 (49), rachis’ R2 >10mm ave. 0 - 0 (- 0.3) (50).  Bracteoles B2 length ave. 27 - 130.3mm (19).  Spikelets on one side of a spike ave. 5.8 - 10.8 (12), length ave. 2.6 - 7.9mm (22), rarely erect usually spreading to patent (47), lanceolate to oblong-cylindrical (26).  Florets on adaxial side variable 1.6 - 7.6 (25).  Glumes ave. 1.4 - 2.4mm long (30), wings lanceolate to acuminate.  Stigma lobes divided <½ to >½ length (36).

Group 2

Stems bluntly trigonous to sharply triquetrous (45). Rays R1 crowded, occasionally 41-50, usually >50 (46),  ave. 112 - 280mm (5) long. Bracts B1 occasionally erect, usually spreading (53), length variable ave. 40 - 88mm (2); prophylls P1 ave. 32 - 45mm long (9).  Spikes ave. 2 - 5 (48-50), ave. length 7.2 - 20.8mm (11); prophylls P2 ave. 1.3 - 6.8mm (21); longest rachis R2 ave. 6 - 54mm (51), sessile rachis' R2 ave. 0.3 - 0.7 (48), rachis' R2 <10mm  ave. 1 - 4.3 (49), rachis' R2 >10mm ave. 0.3 - 2 (50).  Bracteoles B2 length ave. 87 - 161mm (19).  Spikelets on one side of a spike ave. 6.5 - 14.7 (12), length ave. 3.5 - 7.7mm (22), spreading to patent 2 - 3 (47), occasionally ovate, usually lanceolate to narrowly lanceolate (26). Florets on adaxial side 4.2 - 6.6 (25).  Glumes ave. 1.1 - 2mm long (30), wings lanceolate to acuminate (38).  Stigma lobes divided <½ to >½ length (36).

Group 3

Stems shape unknown (45). Rays R1 crowded, generally 4 - 5 (46),  ave. 301 - 380mm (5) long.  Bracts B1 erect (53), length ave. 63mm (2); prophylls P1 ave. 35 - 44mm long (9).  Spikes ave. 4 (48-50), ave. length 3 - 7.2mm (11); prophylls P2 ave. 1.9 - 2mm (21); longest rachis R2 ave. 4 - 47mm (51), sessile rachis' R2 ave. 0 - 1 (48), rachis' R2 <10mm ave. 2 - 3.7 (49), rachis' R2 >10mm ave. 0 - 0.7 (50). Bracteoles B2 length ave. 53 - 114mm (19). Spikelets on one side of a spike ave. 15.7 - 22.7 (12), length ave. 4 - 9.8mm (22), spreading to patent (47), lanceolate to narrowly lanceolate (26).  Florets on adaxial side 4 - 9.2 (25).  Glumes ave. 1.2 - 1.2mm (30), wings lanceolate (38). Stigma lobes divided to ½ length way (36).

Group 4

Stems sharply trigonous (45). Rays R1 crowded, generally 1 - 3 (46), ave. 115 - 150mm (5) long.  Bracts B1 spreading (53), length variable ave. 30 - 72mm (2); prophylls P1 ave. 21 - 26mm long (9).  Spikes ave. 3 - 3 (48-50), ave. length 8.6 - 10.8mm (11); prophylls P2 ave. 0 - 2.3mm (21); longest rachis R2 ave. 3 - 15mm (51), sessile rachis' R2 ave. 0 - 0.3 (48), rachis' R2 <10mm  ave. 1 - 2.3 (49), rachis' R2 >10mm ave. 0 - 1 (50).  Bracteoles B2 length variable, ave. 4.1 - 51.3mm (19). Spikelets on one side of a spike ave. 7 - 10 (12), length ave. 8.7 - 10.6mm (22), spreading (47), narrowly lanceolate to oblong-cylindrical (26).  Florets on adaxial side 6.6 - 8.4 (25). Glumes ave. 1.9 - 2mm (30), wings lanceolate to acuminate 1 - 2 (38).  Stigma lobes divided <½ to ½ length  1 - 2 (36).

Group 5

Stems usually sharply triquetrous (45).  Rays R1 not crowded, generally 11 - 40 (46),  ave. 235 - 330mm (5) long.  Bracts B1 erect (53), length consistent ave. 152 - 165mm (2); prophylls P1 ave. 31 - 45mm long (9).  Spikes ave. 2 (48-50), ave. length 2.7 - 15mm (11); prophylls P2 usually absent (21); longest rachis R2 ave. 2 - 5mm (51), sessile rachis' R2 ave. 1 (48), rachis' R2 <10mm  ave. 1 (49), rachis' R2 >10mm absent (50). Bracteoles B2 length ave. 14 - 109mm (19). Spikelets on one side of a spike ave. 11.3 - 21 (12), length ave. 1.9 - 15.7mm (22), spreading to patent 2 - 3 (47), lanceolate to oblong-cylindrical 3 - 4 (26). Florets on adaxial side 7.8 - 12 (25). Glumes ave. 2.5 - 2.6mm (30), wings absent or acuminate (38). Stigma lobes divided <½ length (36).

Group 6

Stems usually bluntly trigonous (45).  Rays R1 crowded, >40 (46), ave. 250mm (5) long.  Bracts B1 erect (53), length ave. 84mm (2); prophylls P1 ave. 24mm long (9).  Spikes ave. 5 (48-50), ave. length 2mm (11); prophylls P2 absent (21); longest rachis R2 ave. 1mm (51), sessile rachis' R2 ave. 4 (48), rachis' R2 <10mm  ave. 1 (49), rachis' R2 >10mm absent (50).  Bracteoles B2 length 3mm (19).  Spikelets on one side of a spike ave. 7.7 (12), length ave. 29.6mm (22), erect (47), narrowly lanceolate (26).  Florets on adaxial side 9.2 (25). Glumes ave. 4.1mm (30), wings acuminate 2 - 2 (38).  Stigma lobes divided <½ to >½ length  3 - 3 (36).

Cyperus; Cyperus papyrus

Cyperus; Cyperus papyrus